FAMILY Leeuwenhoekiidae Womersley, 1944 [see Grandjean 1973; Diagnostic, Wen 1999:168] [=Thrombidiidae] [?] [=Leeuwenhoekinae ZoologicalRecord, 1963] [misspelling] [=Georgiinae Vercammen-Grandjean, 1973:113] [=Sauracarellini Vercammen-Grandjean, 1968] [of Apoloniinae] [=Whartoniini Vercammen-Grandjean, 1968] SUBFAMILY LeeuwenhoekiidaeIncertae [Brennan & Goff 1977 Jour. Parasit. 63(3):554-566] GENUS Afracarella Vercammen-Grandjean & Kolebinova, 1968:257 TYPE-SPECIES Afracarella africana (Lawrence, 1949) [=Sauracarella africana Lawrence, 1949] GENUS Wagenaaria Brennan, 1967:147 TYPE-SPECIES Wagenaaria similis Brennan, 1967 GENUS Albeckia Vercammen-Grandjeari & Watkins, 1966:74 TYPE-SPECIES Albeckia albecki Vercammen-Grandjeari & Watkins, 1966 GENUS Multisetosa Hsu & Wen, 1963:50 [was subgenus of Sasacarus; Diagnostic, Wen 1984:32] SPECIES Multisetosa acerbus Stekolnikov, 1992:75 [host Chionomys gud (Mammalia); Russia] SPECIES Multisetosa aridicus Kudryashova, 1990:63 [host not given; Kazakhstan] SPECIES Multisetosa asiaticus Kudryashova, 1990:65 [host Allactaga sibirica (Mammalia); Mongolia] SPECIES Multisetosa deserta Kudryashova, 1990:67 [host Meriones libycus erythrourus (Mammalia); Tajikistan] SPECIES Multisetosa horti Kartzev, 1994:94 [host Cricetulus migratorius (Mammalia); Kyrgyzstan] SPECIES Multisetosa hudongensis Ma & Yang, 2002:740 [Parasite of Meriones meridianus (Mammalia), in Chinese, English P. 743; Qinghai] SPECIES Multisetosa lhasae Wen, Wu & Lan, 1984:149 [host Pitymys leucurus (Mammalia); China] TYPE-SPECIES Multisetosa major (Shluger, 1955) [=Leeuwenhoekia major Shluger, 1955] SPECIES Multisetosa montana Kudryashova, 1990:67 [host Cricetulus migratorius (Mammalia); Tajikistan] SPECIES Multisetosa neimongolensis Wen & Yao, 1984:50 [host Rattus norvegicus (Mammalia); China] SPECIES Multisetosa nitens Schluger & Amanguliev, 1972 SPECIES Multisetosa ochotonae Wen & Shao, 1984:173 [host Ochotona pallasi (Mammalia); China] SPECIES Multisetosa persicus (Vercammen Grandjean, Rohde & Mesghali, 1970) [syn. Kudryashova 1990:60] [=Sasacarus persicus Vercammen Grandjean, Rohde & Mesghali, 1970] SPECIES Multisetosa pinguis Schluger & Amanguliev, 1972 SPECIES Multisetosa rybini Kudryashova, 1990:69 [host Meriones libycus erythrourus (Mammalia); Kirgizia] SPECIES Multisetosa safaae Kudryashova, 1990:69 [host Rattus turkestanicus (Mammalia); Tajikistan] SPECIES Multisetosa sciurotamiatis Zhou, Wen, Chen & Zhai, 1987:141 [host Sciurotamias davidianus (Mammalia), English P. 144; China] SPECIES Multisetosa sylvatici Zhou, Wen, Chen & Zhai, 1987:143 [host Apodemus sylvaticus (Mammalia), English P. 140; China] SPECIES Multisetosa xizangensis Wu & Wen, 1984:146 [host Cricetulus kamensis lama (Mammalia); China] GENUS Sasacarus Brennan & Jones, 1959 [Diagnostic, Wen 1981:155] SPECIES Sasacarus incae Goff, 1982:60 [host Lestoros inca (Mammalia); Peru] SPECIES Sasacarus panamense Goff, 1987:105 [host Cryptotis nigrescens (Mammalia); Panama] GENUS Parvithrombium André, 1962:107 TYPE-SPECIES Parvithrombium crassitarsale André, 1962 GENUS Oxithrombium Feider, 1956:846 TYPE-SPECIES Oxithrombium spinosa (Canestrini, 1885) [=Ottonia spinosa Canestrini, 1885] GENUS Morelacarus Vercammen-Grandjean, 1973:323 [for Cordisetoides V.-Grandjean nomen nudum?] TYPE-SPECIES Morelacarus morelensis (Hoffman, 1960) [=Odontacarus morelensis Hoffman, 1960] GENUS Antrohoekia J. de la Cruz & M. Daniel, 1994:72 TYPE-SPECIES Antrohoekia inexpectata J. de la Cruz & M. Daniel, 1994 GENUS Zhongguosha Wen & Xiang, in T. Wen 1984:31 TYPE-SPECIES Zhongguosha banna Wen & Xiang, in T. Wen 1984 SUBFAMILY Leeuwenhoekiinae Womersley, 1944 GENUS Austracarus Lawrence, 1949 TYPE-SPECIES Austracarus procaviae Lawrence, 1949 SPECIES Austracarus campestris Goff, 1990:201 [host Raphicerus campestris (Mammalia); Cape-Province] SPECIES Austracarus cristatus Mahunka, 1984:394 [Tanzania] SPECIES Austracarus lukoschusi Goff, 1983:335 [host Myosorex varius (Mammalia); South-Africa] SPECIES Austracarus masonae Goff, 1983:337 [host Myosorex varius (Mammalia); South-Africa] SPECIES Austracarus reductus Balogh & Balogh, 1983:290 [Queensland] GENUS Austrombicula Lawrence, 1949 TYPE-SPECIES Austrombicula womersleyi (Lawrence, 1949) [=Leeuwenhoekia womersleyi Lawrence, 1949] GENUS Chatia Brennan, 1946 [Diagnostic, Goff & Brennan 1980:836] SUBGENUS Chatia Brennan, 1946 SPECIES Chatia cunninghamae Goff & Brennan, 1980:836 [mammalian host; Washington] TYPE-SPECIES Chatia setosa Brennan, 1946 SUBGENUS Shunsennia Jameson & Toshioka, 1953:89 [Diagnostic, Goff & Brennan 1980:835; to subgen. of Chatia by Traub & Nadchatram 1966:377] [=Parashunsennia Kumada, 1956:257] [Type is harunaensis; syn. Traub & Nadchatram 1966:377] SPECIES Chatia acrichela Wen, Zhou, Chen, Wang & Zhang, 1984:117 [host Rattus confucianus (Mammalia); China] SPECIES Chatia alpina Shao & Wen, 1984:171 [host Alticola stoliczkanus (Mammalia); China] SPECIES Chatia furcoseta (Yang, Zheng & Chen, 1998):270 [Hualong County, Host Apodemus speciosus (Mammalia), English P. 272; Qinghai] [=Shunsennia furcoseta Yang, Zheng & Chen, 1998] SPECIES Chatia harunaensis (Kumada, 1956) [=Parashunsennia harunaensis Kumada, 1956] SPECIES Chatia maoyi Wen & Xiang, in Xiang & Wen 1984:285 [host Rattus confucianus (Mammalia); China] SPECIES Chatia marmota (Wen & Ma, 1997):212 [Taojiang, Host Marmota himalayana (Mammalia); Gansu] [=Shunsennia marmota Wen & Ma, 1997] SPECIES Chatia oudemansi (Schluger, 1955) SPECIES Chatia scabrisetosa (Huang, 1986):277 [host Rattus niviventer (Mammalia), English P. 279; China] [=Shunsennia scabrisetosa Huang, 1986] TYPE-SPECIES Chatia tarsalis (Jameson & Toshioka, 1953) [=Shunsennia tarsalis Jameson & Toshioka, 1953] GENUS Comatacarus Ewing, 1942 [was subgenus of Leeuwenhoekia see Reed 1973] TYPE-SPECIES Comatacarus americanus Ewing, 1942 GENUS Hannemania Oudemans, 1911 [=Batracarus Vercammen-Grandjean, 1968:125] [praeocc. Fain 1961; Type is rouxi] SPECIES Hannemania rouxi Oudemans, 1917 TYPE-SPECIES Hannemania hylodeus (Oudemans, 1910) [=Heterothrombidium hylodeus Oudemans, 1910] SPECIES Hannemania achalai de-Alzuet & Mauri, 1987:114 [host Pleurodema (Amphibia); Argentina] SPECIES Hannemania minor de-Alzuet & Mauri, 1987:112 [host Leptodactylus ocellatus (Amphibia); Argentina] GENUS Hyracarus Lawrence, 1949 TYPE-SPECIES Hyracarus typicus Lawrence, 1949 GENUS Leeuwenhoekia Oudemans, 1911 TYPE-SPECIES Leeuwenhoekia verduni (Oudemans, 1910) [=Heterothrombidium verduni Oudemans, 1910] GENUS Odontacarus Ewing, 1929 SUBGENUS Odontacarus Ewing, 1929 [Redescription, Goff & Loomis 1977:370] [=Acomatacarus Ewing, 1942] [type is arizonensis] SPECIES Odontacarus adelaideae (Womersley, 1944) [=Leeuwenhoekia adelaideae Womersley, 1944] SPECIES Odontacarus apricus Kudryashova, 1976 SPECIES Odontacarus arizonensis Ewing, 1942 SPECIES Odontacarus athertonensis (Womersley, 1945) [=Acomatacarus athertonensis Womersley, 1945] SPECIES Odontacarus attolus (Banks, 1916) [=Rhyncholophus attolus Banks, 1916] SPECIES Odontacarus australiensis (Hirst, 1925) [=Leeuwenhoekia australiensis Hirst, 1925] [=Leeuwenhoekia hirsti Womersley, 1944] SPECIES Odontacarus barrinensis (Womersley, 1945) [Lectotype designated; Southcott 1991:35] [=Acomatacarus barrinensis Womersley, 1945] SPECIES Odontacarus cooki (Southcott, 1957) [=Acomatacarus cooki Southcott, 1957] TYPE-SPECIES Odontacarus dentata (Ewing, 1925) [=Trombicula dentata Ewing, 1925] SPECIES Odontacarus dignus Kudryashova, 1976 SPECIES Odontacarus dracophilis Brown & Goff, 1988:214 [host Draco (Reptilia); Philippines] SPECIES Odontacarus dromus (Womersley, 1939) [Diagnostic, Southcott 1986:173] [=Dromeothrombium dromus Womersley, 1939] SPECIES Odontacarus echidnus (Womersley, 1945) [=Acomatacarus echidnus Womersley, 1945] SPECIES Odontacarus efferus Kudryashova, 1976 SPECIES Odontacarus fedtschenkoi Hushcha & Amanguliev, 1980 SPECIES Odontacarus hushchai Kudryashova, 1995:9 [host Lacerta saxicola (Reptilia); Russia] SPECIES Odontacarus jinanensis Teng-Bi, 1983:75 [host Coluber spinalis (Reptilia), English P. 77; China] SPECIES Odontacarus khanjanii Goff & Saboori, 1998:857 [Hamadan, Asad Abad; Iran] SPECIES Odontacarus langani (Southcott, 1957) [=Acomatacarus langani Southcott, 1957] SPECIES Odontacarus mahaplagensis Brown & Goff, 1988:217 [hosts Draco (Reptilia) & Aves; Philippines] SPECIES Odontacarus mathewi (Southcott, 1957) [=Acomatacarus mathewi Southcott, 1957] SPECIES Odontacarus mccullochi (Womersley, 1944) [=Leeuwenhoekia mccullochi Womersley, 1944] SPECIES Odontacarus mitchelli Goff, 1981:40 [host Rattus niobe (Mammalia); Papua-New-Guinea] SPECIES Odontacarus novaguinea (Womersley, 1944) [=Leeuwenhoekia novaguinea Womersley, 1944] [=Acomatacarus longipes Womersley, 1945] SPECIES Odontacarus patrius (Womersley, 1945) [=Acomatacarus patrius Womersley, 1945] SPECIES Odontacarus pinnatus Goff & Loomis, 1977:372 [reptilian host; Texas] SPECIES Odontacarus saxicolis Schluger, Hushcha & Kudrjaschowa, 1965 [Redescription, Gushcha 1981:41] SPECIES Odontacarus schlugerae Kartzev, 1994:112 [host Gymnodactylus fedtschenkoi (Reptilia); Uzbekistan] SPECIES Odontacarus shandongensis Teng-Bi, 1983:106 [host Gekko swinhoana (Reptilia), English P. 108; China] SPECIES Odontacarus sichuanensis Zhou, Wen, Wang & Zhang, 1981:317 [host Muridae (Mammalia); China] SPECIES Odontacarus southcotti (Womersley, 1944) [=Leeuwenhoekia southcotti Womersley, 1944] SPECIES Odontacarus swani Southcott, 1986:171 [Queensland] SPECIES Odontacarus tetrasetosus Yu & Yang, 1986:64 [host Rattus rattus sladeni (Mammalia), English P. 70; China] SPECIES Odontacarus turkmenicus Kartzev, 1994:114 [host Gymnodactylus fedtschenkoi (Reptilia); Turkmenistan] SPECIES Odontacarus unicus Goff & Brennan, 1978:50 [mammalian host; Venezuela] SPECIES Odontacarus veitchi Southcott, 1986:213 [Queensland] SPECIES Odontacarus xishava Wen & Xiang, in Xiang & Wen 1984:286 [host Rattus confucianus (Mammalia); China] SPECIES Odontacarus xiyi Wu & Wen, 1984:145 [host Sauria (Reptilia); China] SUBGENUS Tarsalacarus Vercammen-Grandjean, 1968:53 [Redescription, Goff & Loomis 1977:372] TYPE-SPECIES Odontacarus bakeri (hoffmann, 1951) [=Acomatacarus bakeri hoffmann, 1951] SUBGENUS Leogonis Vercammen-Grandjean, 1968:53 [as syn.; Domrow 1991:1349] [=Leogonius ZoologicalRecord,] [misspelling] SPECIES Odontacarus adelaideae (Womersley, 1944) [Redescription, Southcott 1989:37] SPECIES Odontacarus athertonensis (Womersley, 1945) [Diagnostic, Southcott 1991:36] SPECIES Odontacarus athertonensis (Womersley, 1945) [Redescription, Goff 1979:144] SPECIES Odontacarus australiensis (Hirst, 1925) [Redescription, Goff 1979:143] TYPE-SPECIES Odontacarus australiensis (Hirst, 1925) [ Redescription, Goff 1979:143] [=Leeuwenhoekia australiensis Hirst, 1925] SPECIES Odontacarus barrinensis (Womersley, 1945) [Redescription, Southcott & Frances 1991:279] SPECIES Odontacarus barrinensis (Womersley, 1945) [Redescription, Southcott 1991:36] SPECIES Odontacarus mccullochi (Womersley, 1944) [Redescription, Goff 1979:147] SPECIES Odontacarus nadchatrami Goff, 1979:149 [Papua-New-Guinea] SPECIES Odontacarus niaoer Wen & Xiang, in Xiang & Wen 1984:288 [avian host; China] SPECIES Odontacarus novaguinea (Womersley, 1944) [Redescription, Goff 1979:149] SPECIES Odontacarus riannae Goff, 1979:150 [Irian-Jaya] SPECIES Odontacarus trisetosus Goff, 1979:153 [Papu-New-Guinea] SPECIES Odontacarus unisetosa Goff, 1979:140 [mammalian host; Papua-New-Guinea] SPECIES Odontacarus ziegleri Goff, 1979:153 [Papu-New-Guinea] SUBGENUS Xenodontacarus Loomis & Goff, 1973:113 [nomen nova] [=Xenacarus Greenberg, 1951:525] [praeocc.?] TYPE-SPECIES Odontacarus maniculatus (Aetna) [=Peromyscus maniculatus Aetna] SUBGENUS Matacarus Vercammen-Grandjean, 1957:625 [1956?] TYPE-SPECIES Odontacarus buretti Vercammen-Grandjean, 1957 SPECIES Odontacarus crimeaensis (Kudryashova, 1995):55 [host Lacerta taurica (Reptilia); Ukraine] [=Matacarus crimeaensis Kudryashova, 1995] SPECIES Odontacarus maroccanus (Taufflieb, 1958) SUBGENUS Orochlorus Vercammen-Grandjean, 1968:122 TYPE-SPECIES Odontacarus micheneri (Greenberg, 1952) [=Acomatacarus micheneri Greenberg, 1952] GENUS Paraguacarus M. L. Goff & J. O. J. Whitaker, 1984:31 TYPE-SPECIES Paraguacarus callosus M. L. Goff & J. O. J. Whitaker, 1984 SPECIES Paraguacarus abrelli Goff & Whitaker, 1984:329 [host Andalgalomys pearsoni (Mammalia); Paraguay] SPECIES Paraguacarus santiagoensis Goff & Webb, 1989:93 [host Octodon degus (Mammalia); Chile] GENUS Whartonia Ewing, 1944 SUBGENUS Whartonia Ewing, 1944 [=Ischnothrombium Feider, 1983] [syn. Daniel & Stekolnikov 2002:150] SPECIES Whartonia delacruzi Daniel & Stekolnikov, 2002:151 [Parasite of Brachyphylla nana (Mammalia); Cuba] SPECIES Whartonia dewitti Nadchatram, 1980:324 [mammalian host; Malaya] SPECIES Whartonia diosi Fain, 2002:69 [Parasite of Megaderma spasma (Mammalia); Philippines] SPECIES Whartonia diploctenum (Feider, 1983):149 [Redescription, Daniel & Stekolnikov 2002:150; La Pluma Cave; Cuba] [=Ischnothrombium diploctenum Feider, 1983] SPECIES Whartonia furcappa Wen & Zhou, in Wen, Zhou & Jiang 2001:195 [Liangshan Yi, Dechang, Parasite of Chiroptera (Mammalia); Sichuan] SPECIES Whartonia kulumadouensis Goff, 1980:494 [Papua-New-Guinea] SPECIES Whartonia multisetosa Goff & Easton, 1991:251 [host Hipposideros pomona (Mammalia); Hong-Kong] SPECIES Whartonia novemsetosa Goff, 1982:376 [host Rousettus aegyptiacus (Mammalia); Tanzania] TYPE-SPECIES Whartonia nudosetosa (Wharton, 1938) [=Hannemania nudosetosa Wharton, 1938] SPECIES Whartonia penthetor Womersley, 1957 [Redescription, Goff 1980:497] SPECIES Whartonia ratnasooriyai Brown, Udagama-Randeniya & Seneviratne, 2003:69 [Parasite of Hipposideros bicolor fulvus (Mammalia); Sri-Lanka] SPECIES Whartonia teongwahi Nadchatram, 1980:327 [mammalian host; Indonesia] SUBGENUS Brennanella Radford, 1954 [subgen. by Vercammen-Grandjean et al. 1965:330; 1945 in ZR 1965; see Brennania Radford 1954?] TYPE-SPECIES Whartonia longispina Radford, 1954 SUBGENUS Asolentria Vercammen-Grandjean, 1968:126 TYPE-SPECIES Whartonia trinidadensis Brennan & Jones, 1960 SUBFAMILY Apoloniinae Wharton, 1947 [Brennan & Goff 1977] GENUS Apolonia Torres & Braga, 1939 TYPE-SPECIES Apolonia tigipioensis Torres & Braga, 1939 GENUS Ceuthothrombium Robaux,Webb & Campbell, 1976 [Thrombidiidae] TYPE-SPECIES Ceuthothrombium cavaticum Robaux,Webb & Campbell, 1976 GENUS Sauracarella Lawrence, 1949 TYPE-SPECIES Sauracarella whartoni Lawrence, 1949 GENUS Straelensia Vercammen-Grandjean & Kolebinova, 1968:253 TYPE-SPECIES Straelensia europaea Vercammen-Grandjean & Kolebinova, 1968 SPECIES Straelensia cynotis Fain & Le-Net, 2000:340 [Parasite of Canis familiaris (Mammalia); France] SPECIES Straelensia tiani Wen, Tian, Guan & Wang, 1996:212 [host Lepus capensis (Mammalia); Shanxi] GENUS Womersia Wharton, 1947 SPECIES Womersia strandtmani Wharton, 1947 SPECIES Womersia midwayensis Goff, Sievert & Sileo, 1989:484 [host Diomedea immutabilis (Aves); Hawaii]